"[A]nimals are not just subject to suffering like man, but subject to much more suffering; their existence is considered to be extremely unhappy, not only because they are exploited and tortured by man but also in nature itself, where the weaker one is threatened and devoured by the stronger, and, moreover, because at least many of them live on disgusting food or in uncomfortable places."
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, "The Early Buddhist Tradition and Ethics", Journal of Buddhist Ethics (1997), ch. 6
https://en.wikiquote.org/wiki/Wild_animal_suffering
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