"During ... a characteristic fermentative yeast flora develops in the liquid... When the sugar is used up and the pH has dropped because of lactic acid production by the lactic acid bacteria, a secondary, oxidative yeast flora develops on the surface of the liquid in the form of a thick, folded layer of yeast."
January 1, 1970
https://en.wikiquote.org/wiki/Fermentation_in_food_processing