"The first scientific studies of Puffins were by on between 1927 and 1939, rather as a sideline to his pioneering work on the . During this period, Skokholm became one of the world's best-known seabird islands. After the Second World War, Lockley turned his attention to and in 1953 published his monograph Puffins in which he described Puffin biology and behaviour in very evocative prose."